Thallichtenberg, Germany
Thallichtenberg, Germany

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Voigt S.,Urweltmuseum GEOSKOP | Lucas S.G.,New Mexico Museum of Natural History and Science | Krainer K.,University of Innsbruck
Palaeogeography, Palaeoclimatology, Palaeoecology | Year: 2013

The Early Permian Robledo Mountains Formation of the Hueco Group in the Robledo Mountains, south-central New Mexico, U.S.A., is a world-class ichnofossil Lagerstätte of supposed peritidal origin. Abundant and diverse invertebrate and vertebrate trace fossils occur in several thin horizons of siliciclastic red beds that are intercalated with thick successions of shallow marine limestone and shale. The depositional environment of the red beds has been discussed for decades but without any detailed sedimentary facies analysis. A recently completed systematic excavation shows that the majority of trace fossils is related to mud-draped surfaces within distal crevasse-splay siltstone to fine-grained sandstone. Interbedded mudstone with well-preserved macrofloral remains and a lungfish aestivation burrow represents a hitherto disregarded facies in overbank fines. Both environmental subzones were at least loosely covered with low-growing plants and experienced common periods of subaerial exposure. None of the physical and biological structures recorded in the measured section indicates tidal influence, but instead they support the reconstruction of a freshwater ecosystem dominated by arthropods and tetrapods. Given their proximity to carbonates of unambiguous subtidal origin, we propose that the trace-fossil-bearing red beds of the Robledo Mountains Formation formed in distal parts of an extensive coastal floodplain during alternating wet and dry conditions. If this interpretation is correct, it provides an impetus to continue ichnological and sedimentological field research in the study area, because the true tidal-flat ichnoguilds still await elucidation. © 2012 Elsevier B.V.


Franz M.,TU Bergakademie Freiberg | Franz M.,University of Gottingen | Kaiser S.I.,Staatliches Museum fur Naturkunde Stuttgart | Fischer J.,Urweltmuseum GEOSKOP | And 7 more authors.
Global and Planetary Change | Year: 2015

The Upper Muschelkalk in the Central European Basin (CEB) is a key example of eustatic and climatic controls on inland seas. The late Anisian rapid transgression from Tethyan waters culminated in a large semi-enclosed inland sea stretching across the CEB. Subsequently, the slow but successive retreat in the early Ladinian resulted in a small remnant sea. The pronounced stratal pattern architectures are translated into a framework of 3rd- and 4th-order T-R sequences. The latest Illyrian 3rd-order maximum flooding surface corresponds to maximum abundances of carbonates and marine phytoplankton. An euryhaline marine ecology is indicated by prasinophycean algae dominating over acritarchs and δ18OP values of 18.9-22.4‰ VSMOW corresponding to Tethyan references. During the 3rd-order regressive phase successive freshening up to hyposaline conditions is indicated by up to 3‰ depleted δ18OP values, shifts to more radiogenic 87Sr/86Sr ratios and maximum abundances of terrestrial palynomorphs. Likewise, 4th-order T-R sequences are constrained by commutated stratal pattern architectures, palynofacies and geochemistry. The favourable correlation of middle Triassic 3rd-order sequences of Tethyan and peri-Tethyan basins demonstrate the principle control of circum-Tethyan eustatic cycles. 4th-order sequences are evident and, although not yet correlatable in detail, indicate 106-year scale eustatic cycles which may be attributed to glacioeustatic sea-level changes. The subordinated control of arid to semiarid low latitude and semihumid to humid temperate mid latitude climates affected the Upper Muschelkalk Sea in particular during 4th-order sea-level lowstands. Substantial fresh water input from Scandinavian sources caused temporal stratification leading to stagnant bottom waters and/or sediments as indicated by palynofacies and U/Th and Ni/Co redox indices. The herein reconstructed middle Triassic zonal climates are in agreement to previously published Late Triassic zonal climates. © 2015 Elsevier B.V.


Klein H.,Saurierwelt Palaontologisches Museum | Niedzwiedzki G.,Uppsala University | Niedzwiedzki G.,University of Warsaw | Voigt S.,Urweltmuseum GEOSKOP | And 5 more authors.
Ichnos:an International Journal of Plant and Animal | Year: 2013

Early Triassic chirotherian footprint assemblages from Poland, Germany, and Morocco are important for understanding archosaur evolution in the aftermath of the Permian-Triassic crisis. However, their ichnotaxonomy is confusing because various authors have interpreted their diversity differently. After an analysis and ichnotaxonomic re-assessment, the presence of the ichnogenera Brachychirotherium, Isochirotherium, and Chirotherium in these assemblages is not supported. Distant similarities with these ichnotaxa are functions of extra morphological variation and substrate-related factors. Instead, Early Triassic chirotherian footprints described under these names are assigned here to the ichnogenus Protochirotherium and to a more slender morphotype identified as Synaptichnium. In particular, Protochirotherium appears to be more widely distributed in central Pangea as a characteristic morphotype reflecting a distinct stage in archosaur evolution. Trackmakers were nonarchosaurian archosauriforms or, alternatively, stem-group crocodylians. Morphologically and temporally these footprints match the hypothetical ancestor of the Chirotherium barthii trackmaker. Chirotherium barthii appears by the beginning of the Middle Triassic. Because of its restricted stratigraphic range, and its wider distribution in central Pangea, Protochirotherium also has biostratigraphic significance for this region and can be considered as an indicator of Early Triassic-aged strata. © 2013 Copyright Taylor and Francis Group, LLC.


Milan J.,Geomuseum Faxe | Klein H.,Saurierwelt Palaontologisches Museum | Voigt S.,Urweltmuseum GEOSKOP | Stemmerik L.,Natural History Museum of Denmark
Bulletin of the Geological Society of Denmark | Year: 2016

A single slab with Late Palaeozoic tetrapod footprints from East Greenland has been housed at the Natural History Museum of Denmark for decades without scientific notice. The specimen comes from the Mesters Vig Formation of northern Scoresby Land in East Greenland and contains a monospecific assemblage of tetrapod footprints that we assign to Limnopus Marsh 1894. As there is no significant morphological difference from other records of this ichnogenus from North America, Europe and North Africa, the described tetrapod footprints can be referred to eryopoid temnospondyl trackmakers. Limnopus is well-known from Upper Carboniferous and Lower Permian continental deposits of palaeoequatorial Pangea. Identification of Limnopus tracks is in agreement with the supposed Late Carboniferous age of the Mesters Vig Formation and thereby also the first evidence of Carboniferous tetrapods from Greenland. © 2016 by Bulletin of the Geological Society of Denmark.


Klein H.,Saurierwelt Palaontologisches Museum | Lucas S.G.,New Mexico Museum of Natural History | Voigt S.,Urweltmuseum GEOSKOP
Ichnos:an International Journal of Plant and Animal | Year: 2015

Procolophonichnium is a globally distributed but rare tetrapod ichnogenus ranging from the ?Late Permian/Early Triassic through the Late Triassic. A Permian age of the holotype from the Karoo of South Africa, as has been proclaimed by some workers, is doubtful. Descriptions lack coordinates of the type locality but suggest instead a position in Lower Triassic strata. Procolophonichnium material from North and South Africa, central Europe, North and South America is re-evaluated. Furthermore, the new ichnospecies Procolophonichnium lockleyi from the Upper Triassic of the Germanic Basin is introduced, based on material with well-preserved trackways that show distinct morphological features, such as an extended “heel” behind digit V. Five ichnospecies are considered as valid: P. nopcsai (type ichnospecies), P. haarmuehlensis, P. nectouxi (new comb.) and P. polonicum as Procolophonichnium tracks from ?Upper Permian, Lower and Middle Triassic strata, and P. lockleyi ichnosp. nov. from the Upper Triassic. Trackmakers of Procolophonichnium were most likely therapsids based on the digit configuration and the phalangeal formula derived from preserved pads. A prococolophonid affinity, as was generally proclaimed, or the attribution to other groups such as captorhinids, is less probable. © 2015, Copyright © Taylor & Francis Group, LLC.


Hminna A.,Chouaïb Doukkali University | Voigt S.,Urweltmuseum GEOSKOP | Klein H.,Saurierwelt Palaontologisches Museum | Saber H.,Chouaïb Doukkali University | And 2 more authors.
Journal of African Earth Sciences | Year: 2013

The Sidi Said Maachou area in the Moroccan western Meseta preserves a succession, up to 400. m thick, of hitherto poorly studied continental Triassic deposits. Recent detailed geological mapping proposes a lithostratigraphic subdivision of the predominantly red-coloured siliciclastic deposits into three formations. Laminated mudstones and fine-grained sandstones in the upper part of the Oued Oum Er Rbiaa Formation have the most interesting fossil content including plant impressions, rhizoliths, fish scales, and invertebrate and vertebrate traces. These biogenic remains are partially associated with tool marks, microbially induced sedimentary structures, oscillation ripples, desiccation cracks, and halite pseudomorphs, suggesting sedimentation in a playa-like, fluvio-lacustrine system under semiarid conditions. All tetrapod footprints from these beds are assigned to Brachychirotherium parvum and indistinguishable from other occurrences of the ichnogenus in Central Europe and North America. Supposed trackmakers are archosaurs of the crocodile stem-group (Crurotarsi) that were widely spread over Triassic Pangaea. Because Brachychirotherium is only known from Late Triassic (Carnian-Rhaetian) deposits, the same age is attributed to the footprint horizon of the Oued Oum Er Rbiaa Formation. This is the first record of Brachychirotherium on the African continent and the first record of Triassic tetrapod footprints in Morocco outside of the High Atlas. © 2012 Elsevier Ltd.


Voigt S.,Urweltmuseum GEOSKOP | Haubold H.,Martin Luther University of Halle Wittenberg
Palaeogeography, Palaeoclimatology, Palaeoecology | Year: 2015

Paleozoic tetrapod footprints are a common and well-known phenomenon in almost all large European countries except for Spain. Here we report on hitherto unpublished vertebrate tracks from Permian red-beds of the south-central Pyrenees that with regard to their relative abundance, diversity and quality of preservation are suitable to fill this gap of knowledge. The described tracks come from two localities in muddy to fine-grained, sandy, alluvial plain deposits in the lower third of the Peranera Formation of the Erill-Castell Basin near Les Iglésies, northern Catalonia. The tracks can be assigned to five ichnogenera, i.e., Batrachichnus Woodworth, 1900, Limnopus Marsh, 1894, Varanopus Moodie, 1929, Hyloidichnus Gilmore, 1927, and Dromopus Marsh, 1894, that we interpret as footprints of temnospondyls, captorhinomorphs, and araeoscelids or similarly sized sauropsids with a lacertoid foot pattern. This ichnofossil assemblage is most similar to ichnofaunas from the Hermit Formation of the Grand Canyon, Arizona, the upper Abo and Robledo Mountains formations of New Mexico, and the main trace fossil site of the Tiddas Basin, Morocco, suggesting a late Early Permian (Artinskian) age for this stratigraphic level of the Peranera Formation. Considering the relative abundance and diversity of captorhinomorph footprints, the new Spanish tracefossil localities may cover the onset of the Early Permian radiation of non-diapsid eureptiles. The thick Late Paleozoic red-bed successions of the south-central Pyrenees have the potential to also bear footprints of otherwise unknown early therapsids, so systematic fossil prospecting of this area is strongly recommended. © 2014 Elsevier B.V.


Hamad A.A.,University of Jordan | Fischer J.,Urweltmuseum GEOSKOP | Voigt S.,Urweltmuseum GEOSKOP | Kerp H.,University of Munster | And 4 more authors.
Journal of Vertebrate Paleontology | Year: 2016

Citation for this article: Abu Hamad, A., J. Fischer, S. Voigt, H. Kerp, J. W. Schneider, and F. Scholze. 2016. First Permian occurrence of the shark egg capsule morphotype Palaeoxyris Brongniart, 1828. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2016.1112290. © 2016 by the Society of Vertebrate Paleontology


Fischer J.,Urweltmuseum GEOSKOP | Schneider J.W.,TU Bergakademie Freiberg | Hodnett J.-P.M.,13008 Arctic Ave | Elliott D.K.,Northern Arizona University | And 5 more authors.
Historical Biology | Year: 2014

δ18OPvalues and87Sr/86Sr ratios were determined on disarticulated xenacanthiform, hybodontid and ctenacanthid shark tooth material from several Early Permian (Sakmarian–Kungurian) continental bone beds of northern Texas and southern Oklahoma as well as from the marine Middle Permian (Roadian) of northern Arizona. The δ18OPvalues derived from the teeth of bone beds are in the range of 17.6–23.5‰ VSMOW, and are mostly depleted in18O by 0.5–5‰ relative to proposed coeval marine δ18OPvalues. This indicates an adaptation to freshwater habitats on the Early Permian coastal plain by several sharks. Distinctly higher δ18OPvalues from two bone beds are attributed to significant evaporative enrichment in18O in flood plain ponds.87Sr/86Sr ratios of around 0.71077 are notably more radiogenic than87Sr/86Sr of contemporaneous seawater. In contrast, the isotopic composition of teeth from the marine Kaibab Formation is characterised by low δ18OPvalues in the range of 13.4–15.6‰ VSMOW while87Sr/86Sr ratios of around 0.70821 are closer to the Roadian seawater value. The distinctly depleted δ18OPvalues cannot be readily explained by fluvially affected freshening in a nearshore marine environment, so a diagenetic alteration of the Kaibab material seems to be more likely, excluding it from further interpretation. © 2013 Taylor & Francis.


Geyer G.,University of Würzburg | Geyer G.,Uppsala University | Peel J.S.,Uppsala University | Streng M.,Uppsala University | And 3 more authors.
Bulletin of Geosciences | Year: 2014

Early Middle Cambrian bituminous coquinoid limestones from a tectonically isolated outcrop in southwestern Kyrgyzstan yield a remarkably diverse fauna, with stem-group cnidarians, trilobites, rhynchonelliformean brachiopods, and other shelly fossils. The fossil site is in the northern foothills of the Turkestan Range and thus forms part of the westernmost extension of the South Tien Shan. The fauna includes two fairly well known trilobite species, Glabrella ventrosa Lermontova, 1940 and Dorypyge richthofeniformis Lermontova, 1940, that provide confident support for an Amgan age of the rocks. New described taxa include the stem-group cnidarian Cambroctoconus kyrgyzstanicus Peel sp. nov., the trilobite Olenoides sagittatus Geyer sp. nov., and the helcionelloid Manasoconus bifrons Peel gen. et sp. nov. Additional fossils within the samples include the trilobites Olenoides sp. A, Kootenia sp., and Pseudoeteraspis? sp.; the rhynchonelliform brachiopods Narynella cf. ferganensis (Andreeva, 1962), Narynella? sp., Austrohedra? sp. nov., and two species of uncertain generic affinity; the tommotiid Tesella sp.; the hyolithelminth Hyolithellus sp.; and the palaeoscolecid Hadimopanella oezgueli Gedik, 1977. Of particular interest is Cambroctoconus kyrgyzstanicus with an octagonal corallum and a sparsely septate calyx.

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